[1] Univ Adelaide, Sch Anim & Vet Sci, North Terrace, Adelaide, SA 5005, Australia
[2] Murdoch Univ, Sch Vet & Life Sci, 90 South St, Murdoch, WA 6150, Australia
[3] INRA, UMR1213, Rech Herbivores, F-63122 St Genes Champanelle, France
[4] Clermont Univ, VetAgro Sup, UMR1213, Rech Herbivores, F-63122 St Genes Champanelle, France