[1] Univ Sains Malaysia, Sch Biol Sci, George Town 11800, Malaysia
[2] Univ Sains Malaysia, Ctr Marine & Coastal Studies CEMACS, George Town 11800, Malaysia
[3] Int Islamic Univ Malaysia, Dept Marine Sci, Kulliyyah Sci, Jalan Sultan Ahmad Shah, Kuantan 25200, Pahang, Malaysia
[4] Kyoto Univ, Grad Sch Informat, Sakyo Ku, Yoshida Honmachi, Kyoto 6068501, Japan
[5] Melaka Dept Fisheries, Batu Berendam 75350, Melaka, Malaysia