[1] Univ Atlantico, Programa Biol, Barranquilla, Colombia
[2] Corp Univ Autonoma Cauca, Fac Ciencias Ambientales & Desarrollo Sostenible, Popayan, Colombia
[3] Univ Estadual Campinas Adao Jose Cardoso, Museu Zool, Campinas, Brazil
[4] Royal Ontario Museum, Ichthyol, Belleville, IL USA
[5] Univ Nacl Colombia, Fac Ciencias, Inst Ciencias Nat, Bogota, Colombia
[6] Pontificia Univ Javeriana, Fac Ciencias, Dept Biol, Lab Ictiol, Bogota, Colombia
[7] Pontificia Univ Catolica Minas Gerais, Programa Posgrad Biol Vertebrados, Belo Horizonte, MG, Brazil
[8] Museu Ciencias Nat PUC Minas, Av Dom Jose Gaspar 290, BR-30535901 Belo Horizonte, MG, Brazil