[1] Chulalongkorn Univ, Fac Sci, Biol Sci Programme, 254 Phayathai Rd, Bangkok 10330, Thailand
[2] Charles Univ Prague, Fac Sci, Dept Bot, Benatska 2, Prague 12800, Czech Republic
[3] Chulalongkorn Univ, Fac Sci, Dept Biol, 254 Phayathai Rd, Bangkok 10330, Thailand
[4] Royal Bot Garden Edinburgh, 20A Inverleith Row, Edinburgh EH3 5LR, Midlothian, Scotland