[1] Sun Yat Sen Univ, Sch Marine Sci, Zhuhai, Peoples R China
[2] Southern Marine Sci & Engn Guangdong Lab Zhuhai, Zhuhai, Peoples R China
[3] Sun Yat Sen Univ, Guangdong Prov Key Lab Marine Resources & Coastal, Sch Marine Sci, Zhuhai, Peoples R China
[4] Chinese Acad Sci, State Key Lab Trop Oceanog, Guangdong Key Lab Ocean Remote Sensing, South China Sea Inst Oceanol, Guangzhou, Peoples R China
[5] Southern Marine Sci & Engn Guangdong Lab Guangzho, Guangzhou, Peoples R China