[1] Chinese Acad Sci, Inst Bot, State Key Lab Systemat & Evolutionary Bot, Beijing 100093, Peoples R China
[2] Univ Chinese Acad Sci, Coll Life Sci, Beijing 100049, Peoples R China
[3] China Acad Chinese Med Sci, Expt Res Ctr, Beijing 100700, Peoples R China
[4] China Acad Chinese Med Sci, Natl Resource Ctr Chinese Mat Med, State Key Lab Breeding Base Dao Di Herbs, Beijing 100700, Peoples R China
[5] Jishou Univ, Coll Biol & Environm Sci, Jishou 416000, Hunan, Peoples R China
[6] Houhe Natl Nat Reserve, Wufeng 443499, Hubei, Peoples R China
[7] Hunan Normal Univ, Coll Life Sci, Changsha 410081, Hunan, Peoples R China