[1] Univ Antwerp, Ecosyst Management Res Grp, Univ Pl 1, B-2610 Antwerp, Belgium
[2] NIOZ Royal Netherlands Inst Sea Res, Dept Estuarine & Delta Syst, POB 140, NL-4400 AC Yerseke, Netherlands
[3] Univ Utrecht, POB 140, NL-4400 AC Yerseke, Netherlands
[4] Univ Utrecht, Fac Geosci, NL-3508 TC Utrecht, Netherlands