[1] Chinese Acad Sci, Inst Vertebrate Paleontol & Paleoanthropol, Key Lab Vertebrate Evolut & Human Origins, Beijing 100044, Peoples R China
[2] CAS Ctr Excellence Life & Paleoenvironm, Beijing 100044, Peoples R China
[3] Univ Chinese Acad Sci, Coll Earth & Planetary Sci, Beijing 100049, Peoples R China
[4] Canadian Museum Nat, Ottawa, ON K1P 6P4, Canada
[5] George Washington Univ, Washington, DC 20052 USA
[6] Fukui Prefectural Univ, Inst Dinosaur Res, Fukui 9101195, Japan
[7] Fukui Prefectural Dinosaur Museum, Fukui 9118601, Japan
[8] Yanji Municipal Bur Land & Resources, Yanji 133001, Peoples R China
[9] Yanji Paleontol Res Ctr, Yanji 133001, Peoples R China
[10] Yanji Dinosaur Museum, Yanji 133001, Peoples R China