[1] Chungnam Natl Univ, Coll Nat Sci, Dept Biol, Taejon 305764, South Korea
[2] Chungnam Natl Univ, Coll Nat Sci, Dept Biochem, Taejon 305764, South Korea
[3] Kyungpook Natl Univ, Coll Nat Sci, Dept Genet Engn, Taegu 701702, South Korea
[4] Yonsei Univ, Coll Engn & Bioprod Res Ctr, Dept Biotechnol, Seoul 120749, South Korea