[1] Univ Nottingham, Sch Biosci, Loughborough LE12 5RD, Leics, England
[2] Aarhus Univ, Dept Agroecol, Blichers Alle 20,POB 50, DK-8830 Tjele, Denmark
[3] Sinodanish Ctr Educ & Res SDC, Niels Jensens Vej 2, DK-8000 Aarhus C, Denmark
[4] Chinese Acad Sci, Key Lab Land Surface Pattern & Simulat, Inst Geog Sci & Nat Resources Res, Beijing 100101, Peoples R China
[5] Chinese Acad Sci, Key Lab Agr Water Resources, Ctr Agr Resources Res, Inst Genet & Dev Biol, 286 Huaizhong Rd, Shijiazhuang 050021, Hebei, Peoples R China
[6] Nat Resources Inst Finland Luke, FI-00790 Helsinki, Finland