[1] Chinese Acad Sci, State Key Lab Systemat & Evolutionary Bot, Inst Bot, Beijing 100093, Peoples R China
[2] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[3] Chinese Acad Sci, Sinoafrican Joint Res Ctr, Wuhan 430074, Peoples R China
[4] CSIRO, Australian Natl Herbarium, Ctr Australian Natl Biodivers Res, GPO Box 1600, Canberra, ACT 2601, Australia