[1] Cent South Univ Forestry & Technol, Coll Forestry, Changsha, Peoples R China
[2] Cent South Univ Forestry & Technol, Key Lab Insect Evolut & Pest Management Higher Ed, Changsha, Peoples R China
[3] Beijing Normal Univ, Coll Life Sci, Beijing, Peoples R China
[4] Mangshan Natl Nat Reserve, Chenzhou, Peoples R China
[5] Univ Ryukyus, Grad Sch Engn & Sci, Nishihara, Okinawa, Japan