[1] Qufu Normal Univ, Sch Geog & Tourism, Rizhao 276826, Peoples R China
[2] Chinese Acad Sci, Nanjing Inst Geol & Palaeontol, Ctr Excellence Life & Paleoenvironm, State Key Lab Palaeobiol & Stratig, Nanjing 210008, Peoples R China
[3] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[4] Russian Acad Sci, AA Borissiak Palaeontol Inst, Moscow 117647, Russia
[5] Nat Hist Museum, Cromwell Rd, London SW7 5BD, England