[1] Penn State Univ, Dept Geosci, University Pk, PA 16802 USA
[2] CONICET UNPSJB, Museo Paleontol Egidio Feruglio, RA-9100 Trelew, Argentina
[3] Univ Nacl Comahue, Inst Invest Biodiversidad & Ambiente INIBIOMA CON, Rio Negro, Argentina
[4] Univ New Hampshire, Dept Earth Sci, Durham, NH 03824 USA
[5] Univ Nacl La Plata, Fac Ciencias Nat & Museo, Ctr Invest Geol CONICET UNLP, RA-1900 La Plata, Argentina