[1] Univ Cocody, UFR Biosci, Lab Hydrobiol, Abidjan 22, Cote Ivoire
[2] Univ Liege, Lab Demog Poissons & Aquaculture, Tihange, Belgium
[3] Musee Royal Afr Cent, Lab Ichtyol, Tervuren, Belgium
[4] Katholieke Univ Leuven, Ecol & Systemat Sect, Louvain, Belgium