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Wellcome Ctr Cell Biol, Inst Cell Biol, Sch Biol Sci, Michael Swann Bldg, Edinburgh EH9 3BF, Midlothian, Scotland Wellcome Ctr Cell Biol, Inst Cell Biol, Sch Biol Sci, Michael Swann Bldg, Edinburgh EH9 3BF, Midlothian, Scotland