[1] Univ Grenoble Alpes UGA, Inst Biosci & Biotechnol Grenoble BIG, Commissariat Energie Atom & Energies Alternat CEA, Lab Physiol Cellulaire & Vegetale,INRA,CNRS,UMR 5, F-38100 Grenoble, France
[2] Univ Manchester, Sch Earth & Environm Sci, Michael Smith Bldg,Oxford Rd, Manchester M13 9PT, Lancs, England