[1] Eastern Univ, Fac Sci, Dept Zool, Chenkalady, Sri Lanka
[2] Wayamba Univ Sri Lanka, Fac Livestock Fisheries & Nutr, Dept Food Sci & Technol, Makandura, Gonawila, Sri Lanka
[3] Univ Peradeniya, Fac Med, Dept Biochem, Peradeniya, Sri Lanka
[4] Tokyo Univ Marine Sci & Technol, Dept Food Sci & Technol, Tokyo, Japan