[1] Univ Rennes, UMR CNRS 6590, Lab ESO, Rennes, France
[2] Univ Rennes, UMR CNRS 6554, Lab LETG, Rennes, France
[3] Ecole Hautes Etud Sante Publ, Rennes, France
[4] Inst Pasteur, Unite Anthropol & Ecol Emergence Malad, Paris, France
[5] CNRS, MNHN, Paris Diderot, Paris, France
[6] ONG Mbou Mon Tour, Nkala, DEM REP CONGO