[1] Univ Queensland, Sch Biomed Sci, St Lucia, Qld 4072, Australia
[2] Murdoch Univ, Coral Bay Res Stn, Coral Bay, WA, Australia
[3] James Cook Univ, Coll Sci & Engn, Townsville, Qld, Australia
[4] Univ Queensland, Sch Math & Phys, Ctr Applicat Nat Resource Math, St Lucia, Qld, Australia
[5] CSIRO Oceans & Atmosphere, Queensland Biosci Precinct, St Lucia, Qld, Australia
[6] Univ Sunshine Coast, Sch Sci & Engn, Fraser Coast Campus, Hervey Bay, Qld, Australia