[1] Chinese Acad Sci, Cold & Arid Reg Environm & Engn Inst, Key Lab Desert & Desertificat, Lanzhou 730000, Peoples R China
[2] Senckenberg Res Inst, D-60325 Frankfurt, Germany
[3] Nat Hist Museum, D-60325 Frankfurt, Germany
[4] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Continental Collis & Plateau Uplift, Beijing 100085, Peoples R China
[5] Lanzhou Univ, Minist Educ, Key Lab Western Chinas Environm Syst, Lanzhou 730000, Peoples R China
[6] Lanzhou Univ, Res Sch Arid Environm & Climate Change, Lanzhou 730000, Peoples R China
[7] Nanjing Normal Univ, Coll Geog Sci, Nanjing 210046, Jiangsu, Peoples R China