[1] Univ Oxford, Dept Infect Dis & Microbiol, John Radcliffe Hosp, Oxford OX3 9DU, Oxon, England
[2] Univ Oxford, Nuffield Dept Pathol, John Radcliffe Hosp, Oxford OX3 9DU, Oxon, England
[3] Univ Oxford, Dept Cardiovasc Med, John Radcliffe Hosp, Oxford OX3 9DU, Oxon, England
[4] HPA Ctr Infect, Resp & System Infect Lab, London, England