[1] Finnish Inst Occupat Hlth, Helsinki, Finland
[2] Linnunmaa Oy, Joensuu, Finland
[3] Inst Mol Med Finland, Digital & Mol Pathol Unit, Helsinki, Finland
[4] Novartis Inst BioMed Res, Basel, Switzerland
[5] Univ Zaragoza, Dept Anat Embryol & Genet, Zaragoza, Spain