[1] Chinese Acad Sci, Chengdu Inst Biol, Chengdu 610041, Sichuan, Peoples R China
[2] Chinese Acad Sci, Grad Univ, Beijing 100049, Peoples R China
[3] Uppsala Univ, Bot Tradgarden, S-75236 Uppsala, Sweden
[4] Chinese Acad Sci, Beijing Inst Bot, Beijing 100093, Peoples R China