Blueberries in commercial production represent several diverse Vaccinium species. Moreover, significant use has been made of the secondary gene pool of Vaccinium, especially in the development of southern highbush cultivars. Section Hemimyrtillus represents species that are part of the tertiary gene pool of Vaccinium. Two species of Section Hemimyrtillus, native to the Portuguese islands of Madeira (V. padifolium Smith), and the Azores (V. cylindraceum Smith) have features of notable value to conventional blueberry development, among these: upright structure, strong growth, abundant flowering and fruiting, good self-fertility, acceptable fruit quality, inflorescence structure suited to mechanical harvesting, and repeat flowering. Among these two species, one is deciduous and one evergreen. Vaccinium arctostaphylos L., a native of the Caucasus region, is closely allied to V. padifolium, hybridizes freely with it, and has many similar characters, and has greater cold hardiness. Initial progress has been made at incorporating these species into forms compatible with V. corymbosum-based blueberry cultivars, and further evaluations are being made of both parental material and the derived hybrids.