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Carthage Univ, IINRGREF LRVENC, BP 10 El Menzah 4, Tunis 1004, Tunisia
Univ Montpellier, LISAH, INRA, IRD,Montpellier SupAgro, F-34060 Montpellier, France Carthage Univ, IINRGREF LRVENC, BP 10 El Menzah 4, Tunis 1004, Tunisia