[1] Univ Tehran, Coll Sci, Sch Biol, Tehran, Iran
[2] Ferdowsi Univ Mashhad, Dept Biol, Fac Sci, Mashhad, Iran
[3] Ferdowsi Univ Mashhad, Inst Appl Zool, Res Dept Zool Innovat, Fac Sci, Mashhad, Iran
[4] Inst Biol Problems North RAS, Dept Biocenol, Magadan, Russia
[5] Univ Free State, Dept Zool & Entomol, Bloemfontein, South Africa
[6] Univ Turku, Zool Museum, Turku, Finland