[1] Univ Nacl Tucuman, Inst Biodiversidad Neotrop, CONICET Natl Council Sci Res, Fac Ciencias Nat, M Lillo 205, RA-4000 San Miguel De Tucuman, Argentina
[2] Inst M Lillo, M Lillo 205, RA-4000 San Miguel De Tucuman, Argentina
[3] Univ Valle, Dept Biol, Grp Invest Entomol, Cali, Colombia
[4] Ctr Invest Sistemas Sostenibles Prod Agr CIPAV, Cali, Colombia
[5] Wildlife Conservat Soc WCS, Programa Colombia, Cali, Colombia