[1] Univ Helsinki, Cent Hosp, Dept Ophthalmol, Helsinki, Finland
[2] Folkhalsan Inst Genet, Helsinki, Finland
[3] Univ Helsinki, Inst Mol Med Finland, Helsinki, Finland
[4] Univ Helsinki, Neurosci Ctr, Helsinki, Finland
[5] Univ Helsinki, Res Programs Unit, Mol Neurol, Helsinki, Finland