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Johns Hopkins Univ, Sch Med, Dept Otolaryngol Head & Neck Surg, Ctr Hearing & Balance,Ctr Sensory Biol,Inst Basic, Baltimore, MD 21205 USA Johns Hopkins Univ, Sch Med, Dept Otolaryngol Head & Neck Surg, Ctr Hearing & Balance,Ctr Sensory Biol,Inst Basic, Baltimore, MD 21205 USA