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Guangdong Acad Sci, Guangdong Prov Key Lab Microbial Culture Collect, State Key Lab Appl Microbiol Southern China, Guangdong Open Lab Appl Microbiol,Guangdong Inst, Guangzhou, Guangdong, Peoples R China Guangdong Acad Sci, Guangdong Prov Key Lab Microbial Culture Collect, State Key Lab Appl Microbiol Southern China, Guangdong Open Lab Appl Microbiol,Guangdong Inst, Guangzhou, Guangdong, Peoples R China