[1] Univ Europeenne Bretagne, ESMISAB, ScInBioS IFR148, EA Lab Univ Biodivers & Ecol Microbienne 3882, F-29280 Plouzane, France
[2] Univ Bretagne Occidentale, F-29280 Plouzane, France
[3] Oceanopolis, Lab Etud Mammiferes Marins, F-29210 Brest 1, France
[4] Univ Europeenne Bretagne, LEMAR, F-29238 Brest 3, France
[5] Univ Bretagne Occidentale, IUEM, F-29238 Brest 3, France
[6] Univ Europeenne Bretagne, EA3880, F-29238 Brest 3, France
[7] Univ Bretagne Occidentale, UFR Sci & Tech, F-29238 Brest 3, France
[8] Univ Europeenne Bretagne, Immunol & Pathol EA2216, F-29285 Brest, France
[9] Univ Europeenne Bretagne, IFR ScInBioS 148, F-29285 Brest, France
[10] Univ Bretagne Occidentale, F-29285 Brest, France
[11] Clin Vet Montaigne, F-29200 Brest, France