[1] Univ Fed Espirito Santo, Dept Ciencias Biol, Lab Genet & Evolucao Mol, BR-29075910 Vitoria, ES, Brazil
[2] Univ Fed Espirito Santo, BR-29075910 Vitoria, ES, Brazil
[3] Univ Fed Parana, Dept Zool, BR-81531980 Curitiba, Parana, Brazil
[4] Univ Fed Oeste Para, Programa Posgrad Biodiversidade, Campus Oriximina,Rodovia 254,257, BR-68270000 Oriximina, PA, Brazil
[5] Ctr Univ Lavras UNILAVRAS, Lab Zool, BR-37203593 Lavras, MG, Brazil
[6] Univ Brasilia, Dept Zool, BR-70910900 Brasilia, DF, Brazil