[1] UCL, UCL Inst Ophthalmol, Div Genet & Epidemiol, London EC1V 9EL, England
[2] Univ Cambridge, Sch Clin Med, Dept Publ Hlth & Primary Care, Inst Publ Hlth, Cambridge, England
[3] Addenbrookes Hosp, Inst Metab Sci, MRC Epidemiol Unit, Cambridge, England
[4] Moorfields Eye Hosp, Natl Biomed Res Ctr Ophthalmol, London, England
[5] Norfolk & Norwich Univ Hosp, Dept Ophthalmol, Norwich, Norfolk, England