[1] Univ Fed Oeste Para, Lab Ecol & Comportamento Anim, Inst Ciencias Educ, Rua Vera Paz S-N, BR-68040255 Santarem, Para, Brazil
[2] Univ Fed Oeste Para, PPGSAQ, CFl, Ave Mendonca Furtado,2946, BR-68040255 Santarem, Para, Brazil
[3] Univ Fed Oeste Para, PGRNA, Ave Men Donca Furtado,2440, BR-68040255 Santarem, Para, Brazil
[4] Univ Fed Rio de Janeiro, Museu Nacl, Setor Herpetol, Dept Vertebrados, Quinta Boa Vista S-N, BR-2094040 Rio De Janeiro, RJ, Brazil