[1] Harper Adams Univ, Dept Crop & Environm Sci, Newport, Gwent, Wales
[2] Smithsonian Trop Res Inst, Panama City, Panama
[3] Univ Western Australia, Sch Anim Biol, Crawley, WA, Australia
[4] CSIRO Ecosyst Sci, Ctr Environm & Life Sci, Floreat, WA, Australia