[1] Cincinnati Childrens Hosp, Div Gastroenterol, Cincinnati, OH USA
[2] Cincinnati Childrens Hosp, Div Dev Biol, Cincinnati, OH USA
[3] Cincinnati Childrens Hosp Med Ctr, Div Gastroenterol Hepatol & Nutr, Cincinnati, OH USA
[4] Kings Coll London, Inst Liver Studies, London, England
[5] Univ Calif San Francisco, Liver Ctr Lab, San Francisco, CA USA