[1] Lund Univ, Dept Anim Ecol, SE-22362 Lund, Sweden
[2] CSIC, Museo Nacl Ciencias Nat, Dept Ecol Evolutiva, E-28006 Madrid, Spain
[3] Univ Michigan, Sch Nat Resources & Environm, Ann Arbor, MI 48109 USA
[4] Univ Michigan, Dept Class Studies, Modern Greek Program, Ann Arbor, MI 48109 USA
[5] Univ Athens, Dept Biol, Sect Anim & Human Physiol, Athens 15784, Greece