[1] Univ Neuchatel, Inst Geol, CH-2009 Neuchatel, Switzerland
[2] Neftex Petr Consultants Ltd, Abingdon OX14 4SA, Oxon, England
[3] Univ Manchester, Sch Earth Atmospher & Environm Sci, N Africa Res Grp, Manchester M13 9PL, Lancs, England
[4] Univ Lausanne, Inst Geol & Palaeontol, CH-1015 Lausanne, Switzerland