[1] Harvard Med Sch, Mass Eye & Ear Infirm, Schepens Eye Res Inst, Boston, MA USA
[2] Harvard Med Sch, Dept Ophthalmol, Boston, MA USA
[3] Univ Exeter, Sch Med, Exeter, Devon, England
[4] Harvard Med Sch, Dept Cell Biol, Boston, MA USA
[5] Peter MacCallum Canc Ctr, Melbourne, Vic, Australia