[1] Kyoto Univ, Grad Sch Agr, Div Appl Biosci, Kyoto 6068502, Japan
[2] Univ Tokyo, Grad Sch Informat Sci & Technol, Tokyo 1538904, Japan
[3] Japan Sci & Technol Agcy, Core Res Evolut Sci & Technol, Tokyo 1538904, Japan
[4] Kyoto Inst Technol, Dept Appl Biol, Kyoto 6168354, Japan
[5] Natl Agr Res Ctr, Tsukuba, Ibaraki 3058666, Japan
[6] Kyoto Univ, Ctr Ecol Res, Shiga 5202113, Japan
[7] Kyoto Univ, Grad Sch Agr, Div Appl Life Sci, Kyoto 6068502, Japan