[1] Univ Montpellier 2, CNRS, Inst Sci Evolut Montpellier, UMR 5554, F-34095 Montpellier 5, France
[2] Univ Cape Town, Dept Environm & Geog Sci, ZA-7701 Rondebosch, South Africa
[3] Univ Free State, Dept Plant Sci, ZA-9300 Bloemfontein, South Africa
[4] Univ Leicester, Dept Geog, Leicester LEI 7RH, Leics, England
[5] South African Natl Biodivers Inst, ZA-0001 Pretoria, South Africa
[6] Univ Pretoria, Schweickerdt Herbarium, Dept Bot, ZA-0002 Pretoria, South Africa
[7] Univ Coimbra, Dept Ciencias Vida, Ctr Funct Ecol, P-3001455 Coimbra, Portugal
[8] Univ Bergen, Dept Archaeol Hist Culture & Relig, N-5020 Bergen, Norway