[1] Chulalongkorn Univ, Fac Grad Sch, Interdept Program Biomed Sci, Bangkok, Thailand
[2] Chulalongkorn Univ, Dept Vet Microbiol, Fac Vet Sci, Bangkok, Thailand
[3] Chulalongkorn Univ, Ctr Excellence Emerging & Reemerging Infect Dis A, Fac Vet Sci, Bangkok, Thailand
[4] Chulalongkorn Univ, Dept Vet Publ Hlth, Fac Vet Sci, Bangkok, Thailand
[5] Chulalongkorn Univ, Dept Vet Med, Avian Hlth Res Unit, Fac Vet Sci, Bangkok, Thailand