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- [3] The late Pleistocene spotted hyena Crocuta crocuta spelaea (Goldfuss 1823) population with its type specimens from the Zoolithen Cave at Gailenreuth (Bavaria, South Germany): a hyena cub raising den of specialised cave bear scavengers in boreal forest environments of Central Europe [J]. HISTORICAL BIOLOGY, 2011, 23 (04) : 335 - 367
- [4] The holotypes of the upper Pleistocene Crocuta crocuta spelaea (Goldfuss, 1823: Hyaenidae) and Panthera leo spelaea (Goldfuss, 1810: Felidae) of the Zoolithen Cave hyena den (South Germany) and their palaeo-ecological interpretation [J]. ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, 2008, 154 (04) : 822 - 831
- [7] Late Pleistocene Crocuta crocuta spelaea (Goldfuss 1823) populations from the Emscher River terrace open air hyena den near Bottrop and other sites in NW Germany: Their bone accumulations along rivers in lowland mammoth steppe environments and scavenging activities on woolly rhinoceros [J]. QUATERNARY INTERNATIONAL, 2012, 276 : 93 - 119