[1] Univ Western Australia, Oceans Inst, Crawley, WA, Australia
[2] Univ Western Australia, Sch Environm Syst Engn, Crawley, WA, Australia
[3] Oregon State Univ, Coll Ocean & Atmospher Sci, Covallis, OR USA
[4] Univ Calif San Diego, Scripps Inst Oceanog, La Jolla, CA 92093 USA