[1] Univ Panama, Herbario PMA, Apdo 3366, Panama City, Panama
[2] Estafeta Univ, Dept Bot, Apdo 3366, Panama City, Panama
[3] Univ Costa Rica, Herbario Luis Fournier Origgi USJ, Escuela Biol, Apdo 11501-2060, San Jose, Costa Rica
[4] Univ Nacl, Lab Ecol Func & Ecosistemas Tropicales LEFET, Fac Ciencias Exactas & Nat, Res Associate,Escuela Ciencias Biol, Heredia, Costa Rica
[5] Ctr Turist Los Chocuacos, Apdo 60303, Puntarenas, Costa Rica
[6] Univ Firenze, Ctr Studi Erbario Trop, Via Pira 4, I-50121 Florence, Italy
[7] Univ Firenze, Dipartimento Biol, Via Pira 4, I-50121 Florence, Italy