[1] Yokohama Natl Univ, Coll Educ, Yokohama 2408501, Japan
[2] Rishiri Town Museum, 136 Rishiri, Rishiri, Hokkaido 0970311, Japan
[3] Nagoya Univ, Grad Sch Sci, Sugashima Marine Biol Lab, 429-63 Sugashima, Toba, Mie 5170004, Japan
[4] Univ Sains Malaysia, Ctr Marine & Coastal Studies, George Town 11800, Malaysia
[5] JAMSTEC, Deep Sea Biodivers Res Grp, Res Inst Global Change, 2-15 Natsushima Cho, Yokosuka 2370061, Japan
[6] Australian Museum, Australian Museum Res Inst, 1 William St, Sydney 2010, Australia
[7] Macquarie Univ, Dept Biol Sci, N Ryde, NSW 2109, Australia