[1] Univ Lodz, Fac Biol & Environm Protect, Dept Invertebrate Zool & Hydrobiol, Banacha 12-16, PL-90237 Lodz, Poland
[2] Univ Helsinki, Ecosyst & Environm Res Programme, POB 65 Viikinkaari 1, Helsinki 00014, Finland
[3] Univ Helsinki, Finnish Museum Nat Hist, POB 17 Pohjoinen Rautatiekatu 13, Helsinki 00014, Finland