[1] Finnish Environm Inst SYKE, Latokartanonkaari 11, Helsinki 00790, Finland
[2] Univ Helsinki, Finnish Nat Hist Museum, POB 17,Pohjoinen Rautatiekatu 13, Helsinki 00014, Finland
[3] Univ Helsinki, Inst Atmospher & Earth Syst Res INAR, Box 27,Latokartanonkaari 7, Helsinki 00014, Finland
[4] Univ Helsinki, Fac Agr & Forestry, POB 27,Latokartanonkaari 7, Helsinki 00014, Finland