[1] Leiden Univ, Ctr Linguist, NL-2311 BE Leiden, Netherlands
[2] Univ Utrecht, Copernicus Inst Sustainable Dev, Environm Sci Grp, NL-3584 CB Utrecht, Netherlands
[3] Max Planck Inst Evolutionary Anthropol, Dept Evolutionary Genet, D-04103 Leipzig, Germany
[4] Univ Groningen, Campus Fryslan, NL-8911 CE Leeuwarden, Netherlands
[5] Univ Wyoming, Dept Anthropol, Laramie, WY 82071 USA
[6] Univ Lyon 1, CNRS, Lab Biometrie & Biol Evolut, UMR 5558, Villeurbanne, France